Otoyo Shrine was built in the Heian period (794-1185) to pray for the healing of Emperor Uda's illness. The Ookuni-sha Shrine, the last shrine, is home to adorable "komane (guardian rats)," and is especially crowded with visitors on New Year's holidays in the year of the rat. In addition to the komane (guardian rats), there are also komainaru (guardian monkeys) at Hiyoshi Shrine, komatobi (guardian flying squirrels) at Atago Shrine, and komainen (guardian snakes) that bring good fortune. The shrine is also known as a place of pilgrimage for its many blessings, such as good health, good fortune, longevity, academic achievement, matchmaking, and safe delivery of children. The shrine is also known for its camellias, which are in full bloom in the spring.